Perhaps the original ECU is more concerned with keeping the catalyst converter happy by maintaining the required 14.65:1 AFR. Also gotta keep the Nox levels low…lose some more performance. It all adds up if the car is emission compliant…
I’ll bet you could tune (MegaSquirt) a 3 cyl GEO metro to get 60 to 65 MPG or more. The emissions would suck though.